JouRNr or TpoprcA! Blor-ocy AxD CoNsrRvAr ro\, J: 5 7 - 65, 200?
Canopy
Structure
of a Logged-over ForestrThirty
YeaNAfter Logging
MOHD ZAKI Han'zahxdd RUSMAWI Mu'la Faculty of Forestry,
Univ ersitr P utra M alays ia, 43400 UPM SERDANG Selanqor, Malatsia
A B S T RAC T- Phytoso€iological studies werc carried out in Comparbnent 128, Pasoh Forest Reserve, Negeri Sembila , in order to assess
the
canopy structureand
vegetation composition of the logged-over dipterccarp forest, thirty years after logging. A total ofren plots, each with the size of 60 x 40 In, werc established
to
assessthe
vegetation composition, species dominance and standstructure
for
the compartment. studies showed that the stand can be categorized as aSrqEiM -
Canaium stand. Meanwhile,the species composition of the study area
car be differentiated into two communities, namely,
Dialiun
kunstleti-
Bacca reasmatrata commnn]'ty and Hopea odonta Palaquium tusttutum co'.mu'nity, and these communities can be turther differentiated into four sub-conununities. The vegetation of the compariment recorded the aveftZe of 12.5%
canopy lay€r r€rovery of the emergent layer (SD, znd 40%, 4l .5Eo,25 5%, and 17 5% lor
the
dominant and co-dominant (T1).understorcy (T2), shrub (S) and herb (H)
layers, rcspectively. The family with the most representative was Dipterocaryaceae with twelve species representing three genera, followed by Leguminosae.
tr JMrdr:
c
opy stlucrur€ recove.y, logged-overINTRODUCTION
Peninsular Malaysia is blessed wift a unique tropical forest ecosystem. The forest, which is a highly valuable e€onomic rcsource, has be€n systematically managed by the Forestry Department Peninsular Malaysia since
lml
.For instance, lhe lowland and hill dipterocarp forests have been managed under ihe Malayan
Uniform Systen
(MUS) and Selective Management System (SMS), respectively. Even though the forest is being hanested annually, harvesting is carded out according to the strict guidelines of MUS and sMs.The MUS was
first
formulated into the approved system at the 1949 Colferencq of Malayan State Forest Officen (Wyatt-Smith, 1995), and was first implemented in 1955(Yusuferat, 1986). According to dle guideline of the system, all mature arees of commercial species above 45 cm
in
DBH are to be removed in a single harvesting.Naturally. the forest
will
recover afterha
esting, aDd in the case of rhe MUS, the forest is expected to recover lvithin 50 to 55 years (wyatt-smith, 1995). However rhe lateof th.
d.mageduing harve*iog. an N.eqi Sdbilo,
b*ssdeluva-y
sbtus or dE losg.d-ovtrrorrsd rlmr
whicr,ws
hftn.d h
190s uddd rh. MUs.MXE'IAIJANDIIEItrOE
The sody
w6
cobducred incompdnei.
123 of?6oh
FosrRsft,
Neeais.mbild
(Fig. D,
a.ddins
b {re upubush€d Ecordor
lneForc$ry D?pa
menrof Nes.i smbilan'lhe.ompml9afiflhNstd
loidrMd.Fi.Meb.losicds iqrhe
strdy
!rc.
intei.r.l reeie4 fthtively
nodffiEsvdF mud
nidtur or r 304nn
(lem h De7). rtesiE is cralsir,ed a
o 3d
whjcn
sciv.r
havr ftinrarr (wer mmrh) inNovmb
ad De4bbd(Mn,
1992).Thelsde]' Assstmiof conpo]fu,23 ftcdv'y
Guccssiotr)ol
rhelogg.d-ov'
vcgehlion$udiesb,sedmlh.fulhodol 3Eu
Blaqud€ujivm, l$7;
Mobd znri.l99rMohdz3ridrar, zoori zmo
'fttried
phybsociolosic,l
rtl.vrs
or srmple ptob (each*ith
thc size of 60 x a0 m) wer.sbbrsh.dmdodny in
rrt.ooparffi
The phyrosociolosicard.v6s
.otrld be rimptyC{NoN
snoauE
oF^irffDovs FBr r) S.l.dioi of
sites*irhonr
crps mdb) rd.idrdioi
dEord
of 3ll spe.nslmd
in.achot$env. by6: dcrged
or supd
@ (sT
rbov. 30n
in@l
h.icnq
domimt Ol: ls 30b),c0'
dom,nei,jduddbEJ
crl: 3n),
suppEs.n mdstrb
(s: z-
3 m)0dherb (Ir:
b*.rrhs
2b).
c) E
inarion or crnopy coYeBsc md sociabilit lorerh
speies jnab lryd (bsd ontuiivd
o937).rd
d) Id.nrilication
or phyro3ociotogicalRESIJIISANDDISCUSSION
Th?
eddgd
layu (sj[) ofconptu.
r23whichwshds@dh 194( 4)iam.h.d a
almge hejght of 36ft,
w b 6ve Erevts Ed'd.nd.heisntol40nvhil.rh.lov*r s.
llyd
ws Mojd.d by R.re*s r. 2 ad 6 (eschwilnteelheishof
onrGable ulb€Esutb rlso sbow rlar dEw@.
h.i!h! of llETl ]!ydu7Jm,*h
elne IEmh.ichrofdE T2.S
edsw
16B7.1m?dl.654EpatiEly A! lhom
in Trble r, n\.ebd
ing4qat
srbid.d low
b nodmE sr-hyd
cov.Es.of596 b 20%, *nh
d.
avdgc of r2.59r. oD$.
orhdhan4hecmp!@it
hdnodd
b high.oven€eof'I md T2
hy6,
witn rh!sp{rively.
Th. @opy-tay{orcnge
otrhc s-laya8
ms.d
fiDm 20* b 30*. dd[E
b
didrt ro ducidar rbe rcove'ys@s
ofrbe
!s4
rh. heishr rcovery md.eopy,
fis@ t
Loatioo ofC.Eldftd
123, IhlobF6s. RM.,
NesdiS.EbiL!
Shlt
ofNegsi Sdtild
Compt l2a
to L\e
d
' oln\epnm
yfo6t(PD, *bilh
*cE
obrlinedod Epdcd
prcviously by MoMzli
(1eee). rheimhs
sbor dar thea€n8eheiotof
ssYr-hFof Comptua.
r23 (F74)
'.lbturicdly
lowrhm
&. rv.nse l'.ighloflh.PFof4sn'Tbasamcmb€saidroih. ms
sT-lrydcoEns. olrhaFT4 sibsofl,,5{jwbichissig.jJimdyl*dthmlh.
PF (nem (:()r€@ 01269b) (riblc2l.T abb 2
a'dT2iscoDplablelolhatolth.PF,
hovev{. thecov.rag.
or6.
S mdH nb*d.
Accoldi4btbcddislical,Iy.is'lh.t
we'e sisnificant difieEocer
belv.d
thccov.ng.-pft.dbc.
ofTl hy.r
of rhe 10significd
dincHt l0
6colts fu hyd
Sp!.iB
ri.nnd
rndconlotilid
A
bbl
of 101 sp.cierpasddns
?3ea
relev€s. The
rcrdiv.
nchnes or g.nos ro fthnss olfanily to gen$ dchn*s wa55*
Cr'bb 3).Thescngs.
econpdabl.
bln.
dudies conductn on fto
ndd! h@6td ii
r973inS.d,InC InaFoBiRffiq N.sdi
Tn. Y.g.rtjdn or
d. cohp'fumt
cm be(lJ2)si!r - ca,,.i!n
and). anc mos comod 6peci6ii
neonplrmut bdidd Syljim
sp. andC@rrl4
sp.@
Pdrr;aatutd obtrtilotir
md
ri.ura r,ir\,#,
wNche EpEsnrd
in nine ou.
ol
Gn Elev6slTabt
4), Tn.xdEq\.dd|aid.Pdk{lin6tfun'
Elw6.!stelevalionro3i4nlm200b260n
CaNory
sul
E oF ALmED{@ m6r
abov.
!a l.v.l (r s
L).Aroigd
rh.@ii speiesinn\iscommuiymDt!dl,,','
.
amtu\
Mdtt b
s9., 5 horc arer qb
roc(tlrllMtPai*of thecol]llNit
such d H,.dotutq
Pak46m tuhsdti,
PretM,
DipktocotpB
zraditotu: bd sinto
h gcn.i,l.
$a. sh&
hrv.novdtd
rrinyv.ll
wiln low ronod.nt STlay.i.
md(s@prd6o.
lrsi hoi ,
r99D.d.
highrcpresented
fanily vith 12
speciesindividulsineehofdElopRolay$is
appenrly
.rall,
TIE ST andrl
hyes.fu
sp''ct.kil'p'suphimwfupodI]
dipt.rocdpr
suchftlvious
rcpm
by Mobdz*i
(1eee)hd
Mohd
zlt
e''r.
(2004)sbrd
dlar mo$ orrhe
!b& hd.*.d odd nr
MUsbd
ro(miiry.mnipM.rpt
ifrnedirdy,f br hdEslin3'Ttephemmmisdand.d ini\epEssfudy'Tbaoory*pldadonfol
the lack ol dipcmcdps in $e sbds mly lie
in $o
bril nMbd
ofdiD€r@rDs Eken ourMOIDZ^Igl{hziARUSMAVTMd.
fthb
ri Heighr (n) udcovds. Pmdbse{*)ln4b
cropytrydfqdi6@!!r.ldf
-E;i:iffi---i-- - -i----l
r$h2: cmloismof sds.ft 6r,rrrqffi &hdreo
F 4adbn'ryf.N
Dbh3 codponu orh{m. o.hsb.kd rmddy ro
No.oIO.s, NoolPuili6 Frorsir(brj
No. olsFci6
?ft.nunhshpdanhaBi i@rntusiddiErothdofliEErrlo**bmdic
re!
rdio of g.dusb :p€i$
n.hna3(8Eu olom) dd
ndo offmily b ghur
trhle,l: slljfr&yof $. cll|lllM lpets,nd
difitrDriallpecidoib..dmuiri.N
,i
'i .i
cd&tFj6ier,'sftlh,d,
i sa6r
ryrd (tu tojrqu. Dn;
ol nre
$id dDig bNBrins.
TheEords ol
Forcsl-y Depment or Negedscnbils tqice, in
1960s enddipr.rccarps. which
my
hrveEeb.n fi.
DBH ol
noE
rhin 45cn
@urdMv. b.d
bretr
ornEdpbrRTs
vhichecnffidy
for tlEn ro
.frs.
md donimE ihe shndr.eely pb,a ora
dddl+$.esion
Fnoddu
succe$ioilr
3p.di.si sn.h as,.Jd.dtu baccat!\ Eadospdnr
diadenunatd Nn |MnLto a.,ia't1E opffiis
drtg
ea.fisohd*bhsdmmc.dlhse
domind. n\e
s
(h. siDce rhenunbr
md6vds.ordhtscrJpsmElarirclysm,
fiese
sbds
could be plsnred with soncThe follovilc
can
be mrde lron
r. rbe fo;* io
rhc6dpdrnm, hs r.oved
fr0ywll
(ow bnodtu)
inshon pcnod
orhr.
(ahnt rhiiyY.dt.
tha mn diprmcaps
$iny yat artr
hNed. Thes. 3bnds EquiE repldring of sone
rpc.ts fmn
lne diptemcllpbe nsde on rbe
dlb
lrcm tbe ves.h6oistudies, pEeios losging bd
pc.f.,,tic inve ori* ro.lDcidft
rbe d.grcd of.l6en$ ordiflmic
of$s rccovdngA..oa. lt99t, Fone
Resaurc. M@P^i
rtdt Mdldytu.(e .d.)sD
.nt
olgmiedo,UnibdNadmDeyelopnd PFgm (F.Ao{-NDP}Krd.ru9E
^non tteei. ashcttui
M4!Pai8ukr M4rar,4.
(46 ed.) Narionar MappingD?p|,M,ray!;qKu,lrlu'pu
Amri.nj,
s, (z{xrz).cohpaiM of@v.Jt
s@s oftwo lossln-oYd
.@
in N?seiS.nbilsn. ,. s..
"r,JB, Frc
ry orFqiturn, K,oear).Aim
d metho& or phyrosociolos or "rgebriod$nic
.Pp. 60?-623. Id: Anod (.d.). PUFB
,n
nmoD ol Dr sotodli Notnhhi. Ko6.Mohd
zrki.
H,(1999).FodEovery
d rehabiharjoiNso*hcns jn Ncgdi
Sdbihn, Malaysia, bi$d onv.g.dion Uiivdiry,
r,pm (upublished).r&*dzr'4H,LnH0 rr]{-Mdtur,i.a64 r'l
MI(Minrm[
M.K YNor& L [\iiwr
(2{ol), RENery ol lwlanddDbom+ fo(s! !nd6
th.Md'yd Uniforn Sy$.m, topld So.iary.l
rmpi.,r Ecology,
n
p,iqlall)l0l ll0.
Motd Zali, H., A,Arilin, S.
Nclnbdg,A,
l!r.hmdAa.!LEIrr@laLlb.n
&lnNi*Munlnd,
(r[06),cm'pqftm ofEFbdd omsirion
inchlbilibtn
,€gds.md!4fof$inlsohFm$
MOIIDZAKIH
ZN&RUSMAWIMUd'S..! bo!
E, (1973).Indlddion b rht M.la$ib r L
P.:'al[6is diDs3.
Mdldt NtuEta!tuI24\2)ttt tu.
ry.lr-$drn,
J.O99.
Ntanu,lof Mslaymsirvi.utft
fd.r,ind
FoBts. (2.d).
Matay&
Fte R@dt No.z3.Fdat ReFlrI Ldjft Mrbrli4 x.F
'e.
ltd,E rr,A!1,4[r.sriiat&A,Kd'
(1936). rnplicarion
of
rhcsot*live M'rr8!@r Slrtu
otrre'mrsd{t
of
rb.binrlEb offtDi d6M.r.yria.
l!.6t30.
rn:Ylst
rI,A.xmjl
M, NikMuhasad & M. shuki (edt.
Pne.diry,
otw.rr:^q .n hard
oJMatr Activiti.! .n T.aiicat uphna
Fozdr.@$ni. u,ivdiii P.@bn
M,irysi4s.dm8,
r.6 F.bruary 1934Uni6iliP!fuimM,lrygj*